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I've heard/read that Campagnolo horizontal dropouts are a carryover from their early Cambia Corsa derailleur geared dropouts, which allowed the axle to roll forward and back to slacken the chain and allow shifting under movement.

A Campagnolo masterpiece: the Cambio Corsa – B.G. legendary Bikes (bglegendarybikes.com)
And it worked! Assume that's a free wheel.

The dropouts look just like their later ones, minus the adjusting screws. Investment cast, man. Rock hard. Clamp that wheel in there with Campy skewers, and that thing won't come out!
 
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