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Are you racing? Cause if you are, you should get some solid tubular wheels with either a challenge or tufo tire, depending on where you live. Ultegra hubs/ Mavic Reflex rims, or some such.

Often you can find DA hubs on tubular wheels for the same price or cheaper than Ultegra hubed wheels. The DA hubs have better seals, IMHO. But wunlap is spot-on about wheel choice. Get one that is rebuildable immediately. Tubies will also be the single most beneficial upgrade for racing you can make.

About forks ... If you like the "old school" steel fork look, check out the Wound Up. It's got straight legs but it looks pretty cool. If performance is what floats your boat, get an Easton carbon fork. You will notice a difference hefting the bike by replacing your Al fork with a carbon.

 

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I'd get tubbies before I changed my fork. I'd add, are your contact points to your liking? Do you like your saddle? Your pedals? handlebars? I'd change any of these things first, then get tubbies, then a carbon fork.

Pedals are a big deal. ATAC's or Eggs/Candies (I prefer Eggs). I'd think bars would have to be way narrow for me to want to swap but then I like the bars I've got and might be singing a different toon if they didn't suit me. I've been running an ancient Flite on my race bike for a while. The seat needs to be sturdy but not necessarily comfy for long rides. Still, stock saddles are often crap ...

Carbon fork should be the final upgrade for sure.