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Kinda sounds like you maybe just overtaxed the knees with too little extension due to a low saddle height to me. This can happened to me w/ an optimal setup if I hammer a bit too much on sprints and/or climbs on a longish ride or spin above 115-120rpm for a long period of time as well.

I'd say move the saddle back to its previous height and try to do the 62 mile route at an easy pace. If the knees are starting to feel significantly worse around 20 miles in, head back in and make sure your fit is good from a KOPS and leg angle at full extension perspective. Not sure what options you have w/ Speedplay, but moving the cleat slightly more towards the arch side of the ball of the foot seemed to help me a bit as well.

.02
 
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